Civilians have borne the brunt of unlawful killings, kidnappings and the widespread destruction of houses during the four-year separatist conflict in Cameroon, Amnesty International says.

The rights group collected witness accounts and analysed satellite images to study the impact of the violence.

In a series of attacks in February this year more than 4,000 people were forced from their homes in Nwa in the North-West region.

They were targeted by Fulani vigilantes who are accused of siding with Cameroonian government forces.

Amnesty also found evidence of reprisals throughout the conflict by English-speaking separatists against Mbororo Fulani communities which had left more than 160 civilians dead.

The report concluded that all parties to the conflict had committed human rights violations and abuses whilst civilians had been caught in the middle.